In the following passage, some words have been deleted. Read the passage carefully and select the most appropriate option to fill in each blank.
The earliest kind of print technology was developed in China. This was a system of hand (1). From AD 594 onwards, (2)_ in China were printed by (3)______ paper, against the inked surface (4)______ woodblocks. As both sides of (5)______ thin, porous sheet could not be printed, the traditional Chinese ‘accordion book’ was folded and stitched at the side.

Select the most appropriate option for blank number 4.

1. A
   of
2. B
   in
3. C
   by
4. D
   from