'P % Q' means 'P is the brother of Q''P + Q' means 'P is the daughter of Q'.'P = Q' means 'P is the son of Q'.'P # Q' means 'P is the wife of Q'.If L % M + N # R = S, how is L related to S?

Let's break down the expression L % M + N # R = S step-by-step:L % M $\to$ L is the brother of M (${\text{L}}^{(+)}-\text{M}$)M + N $\to$ M is the daughter of N (${\text{M}}^{(-)}\to \text{N}$)(So L and M are siblings, and N is their mother/father)N # R $\to$ N is the wife of R (${\text{N}}^{(-)}\iff {\text{R}}^{(+)}$). This means R is the father of L and M.R = S $\to$ R is the son of S (${\text{R}}^{(+)}\to \text{S}$)Now let's determine how L is related to S:R is the son of S.L is the son of R.Therefore, L is the son of the son of S, which means L is S's son's son (grandson).